Suffering is a state of mind that is difficult to measure and analyse in human beings and considerably more so in animals. It is related to the environment in which we live and our physical and mental states. Understanding the physiology of suffering in animals is crucial in assessing animal welfare. Written by an expert in applied welfare aspects of physiology, this book is the first to address the physiological aspects of suffering in animals. It explores the different causes of suffering – physical discomfort, thirst and hunger, the responses in the body that lead to suffering and it offers insight into how suffering can be managed.